(4) Water limits and fire Explanation: Grass ecosystems are an early stage of succession in regions where the mature ecosystems are forests. However, grass ecosystems are climax ecosystems in grassland regions, where there is not enough rainfall to support a forest. Due to limiting, factors such as water scarcity and forest fire, a grass land ecosystem changes to a desert ecosystem. The ecological succession of grassland is desert. It is, however, notable that the soil of grassland ecosystem is rich in nutrients and organic matters and provides the largest biomass.